LOS ANGELES -- After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

SAN DIEGO — San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by far the zoo's oldest resident, in what has been a year marked ...

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of ...

A few years ago, we added a tortoise to our legion of pets at home, and it's basically like having a tiny dinosaur as a roommate. There was a bit of a learning curve at first, as a family who had only ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

Add Yahoo as a preferred source to see more of our stories on Google. LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved ...

A zoo visitor greets Gramma in 2015 at the San Diego Zoo when the tortoise was a spry 130-year-old. (Mark Ralston / AFP via Getty Images) Gramma, a Galápagos tortoise whose life at the San Diego Zoo ...

SAN DIEGO — One of the first animals brought to the San Diego Zoo has died at the age of 141 years old. The zoo announced the death of its beloved Galápagos tortoise, Gramma, on social media, saying ...